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Two men arrested in the death of Cecil the lion — a case in which an American dentist has also been accused, unleashing a torrent of anger online — were released Wednesday by a court in Zimbabwe on $1,000 bail each.

Theo Bronchorst, a professional hunter, and Honest Trymore Ndlovu, a land owner, both Zimbabweans, said through their attorney that they were innocent of poaching charges, which officials said could bring a sentence of 10 years in prison.

Zimbabwean authorities said that Walter J. Palmer, a dentist from Minnesota, paid at least $50,000 for the hunt. Palmer has said he relied on the expertise of local guides “to ensure a legal hunt.”

But the lion that he and his local guides killed wasn’t just any lion, according to Zimbabwean officials.

He was Cecil, a major tourist draw at Zimbabwe’s Hwange National Park.

The 13-year-old lion, recognizable by the black streaks in his mane, suffered a slow death, according to the Zimbabwe Conservation Task Force.

The hunters lured him out of the sanctuary of the park with a dead animal on top of a vehicle, the conservation group said.

Palmer, officials said, then shot the lion with a crossbow, a method for which he is known. But Cecil survived another 40 hours until the hunters tracked him down and shot him with a gun.

Cecil was skinned and beheaded, and the hunters tried to destroy the GPS collar that Cecil was wearing as part of research backed by Oxford University, the group said.

“I had no idea that the lion I took was a known, local favorite, was collared and part of a study until the end of the hunt,” Palmer said Tuesday in a statement. “I relied on the expertise of my local professional guides to ensure a legal hunt.”

Torrent of anger online

His alleged role in Cecil’s death brought a wave of online anger crashing down on him.

The Yelp page for his dental practice in Bloomington, Minnesota, was inundated with reviews posted by people irate over his lion hunting.

“Shame on you, killing a majestic creature,” wrote a user named Charmie P.

The website for Palmer’s business, River Bluff Dental, appeared to have been taken down.

A torrent of outrage flowed on social media, with celebrities such as Sharon Osbourne lambasting the dentist.

“I hope that #WalterPalmer loses his home, his practice & his money,” Osbourne tweeted. “He has already lost his soul.”

At least $50,000 allegedly paid for hunt

Investigations suggest the killing of Cecil was illegal because the land owner “was not allocated a lion on his hunting quota for 2015,” said a statement from the Zimbabwe Parks and Wildlife Management Authority and Safari Operators Association of Zimbabwe.

The dentist said in his statement that no authorities in Zimbabwe or the United States had contacted him but that he would assist them in any inquiries.

“I deeply regret that my pursuit of an activity I love and practice responsibly and legally resulted in the taking of this lion,” Palmer said.

But Cecil’s killing doesn’t appear to be the first time Palmer has got into trouble while hunting.

A man by the same name and age, and from the same town, illegally killed a black bear in Wisconsin several years ago, according to court documents.

That individual pleaded guilty to making false statements knowingly to the U.S. Fish and Wildlife Service, and was sentenced to one year on probation and ordered to pay a fine of nearly $3,000, records show.

A New York Times article in 2009 that profiled Palmer and his hunting methods said he had served a year of probation over the false statements case.